Graduate Degree Program Summary

Graduate degrees offered


  • M.S.; Ph.D.


A specialization is a well-defined area of study that will appear on your transcript with your degree and major. Specializations are optional in most UNL programs.

  • Bioinformatics
Areas of Study
  • Biophysical Chemistry
  • Biotechnology
  • Metabolic Biochemistry
  • Molecular Genetics
  • Molecular Medicine
  • Plant Biochemistry and Molecular Biology
  • Structural Biology
Promo image for Biochemistry

Contacts for Biochemistry

On the Web


Graduate Chair

Dr. Mark Wilson

Graduate Admissions Co-Chair

Dr. Jaekwon Lee


Graduate Admissions Co-Chair

Joe Baryicki


Campus Address

N200 Beadle Center

Lincoln NE 68588-0664

Application checklist and deadlines

Required by the Office of Graduate Studies

See also: steps to admission.

Required by Biochemistry in GAMES

After you apply, allow one business day for us to establish your access to GAMES, where you'll complete these departmental requirements:

  • Entrance exam(s): GRE (Biochemistry subject test recommended)
  • Minimum English proficiency: Paper TOEFL 550, Internet TOEFL 79, IELTS 6.5
  • Three recommendation letters
  • Curriculum Vitae or Resume
  • List of published papers, if applicable
  • Statement of purpose

When sending GRE or TOEFL scores, UNL's institution code is 6877 and a department code is not needed.

Application Deadlines for Biochemistry
Fall: December 15 Spring: September 1


The Center for Biological Chemistry (CBC) offers a graduate program which is interdisciplinary. Research opportunities exist in a variety of areas, including plant biochemistry, photosynthesis research, plant and animal metabolism, metallobiochemistry, enzyme mechanisms, cancer biology, redox biology, and molecular biology. Biochemistry faculty and participating faculty in animal science, agronomy, chemistry, and biological sciences contribute to the program.

Instructional programs are focused in laboratories and classrooms within the George W. Beadle Center for Genetics and Biomaterials Research. The Beadle Center houses most of the department-affiliated faculty members in state-of-the-art research laboratories and many of the Center for Biotechnology and Redox Biology Center core facilities. These facilities include mass spectrometry, spectroscopy, X-ray crystallography, plant transformation, and microscopy facilities. Also available are an outstanding full-service plant growth and greenhouse complex, and equipment for fluorescence-activated cell analysis, surface plasmon resonance, analytical ultracentrifugation and next-generation high-throughput DNA sequencing.

Courses and More

Students in Biochemistry are most likely to take courses in: See also: Course Catalog in the Graduate Bulletin.

Students will work with an advisor to create a Program of Studies or Memorandum of Courses during the first half of their coursework.

Faculty and research

Jiri Adamec
Oxidative Stress; Proteomics; Metabolomics; Cancer; Neurodegenerative Diseases; Predictive Biomarkers
Joseph Barycki
Crystallographic and Biochemical Characterizations of Enzymes
Donald Becker
Redox Enzymology; Gene Regulation; Multifunctional Proteins; Proline Metabolism
Paul Black
Fatty Acid Transport; Fatty Acid Transport Proteins; Metabolic Transformations
Nicole Buan Murphy
Energy Conservation; Redox Biochemistry; Microbial Physiology
Edgar Cahoon
Functional Genomics; Sphingolipid Metabolism and Function; Nutritional Biofortification of Crop Plants
Concetta DiRusso
Nutrient Fatty Acids; Fatty Acid-Dependent Regulation; Fatty Acids in Health and Disease
Mark Griep
Biodefense Research; Antibiotic Discovery; Pathogenic Bacteria; DNA Replication; Bioinformatics
Edward Harris
Stabilin Receptors; Glycosaminoglycan Metabolism; Receptor-Mediated Endocytosis
Tomas Helikar
Computational Modeling of Dynamics in Large-Scale Biological/Biochemical Networks; Host-Pathogen Networks; Development of Technologies to Enable an Integrated Systems Biology Approach in Research and the Classroom
Oleh Khalimonchuk
Redox Sensitive Signaling Systems
Jaekwon Lee
Metal-Ion Metabolism; Copper Transporters; Oxidative Stress Response
Jonathan Markham
Sphingolipid Metabolism in Plants and Algae
John Markwell
Photosynthetic Efficiency; Higher Plant Photosynthetic Pigments; Soybean Proteins as Allergens
Kenneth Nickerson
Fungal Dimorphism; Quorum Sensing in Fungi; Bacterial Detergent Resistance; Microbial Insecticides
Lawrence Parkhurst
Rapid Reaction Kinetics; Laser Spectroscopy; Eukaryotic Transcription
Seung-Hyun Ro
Adipocyte Biology; Autophagy/Mitophagy; Mitochondria Metabolism; Oxidative Stress; Sestrins/mTOR Signaling
Rebecca Roston
Role of Membrane Dynamics in the Plant Cell using Combination of Genetics, Molecular Biology, Protein Biochemistry, and Biophysical Approaches; Membrane Response to Freezing Stress; Understanding Molecular Mechanisms of Photosynthetic Membrane Biogenesis
Gautam Sarath
Biomass and Forage Qualities; Warm-Season Prairie Grasses; Prairie Legumes; Seed Germination and Plant Development; Histone Biotinylation
Melanie Simpson
Prostate Cancer; Tumorigenesis Mechanisms; Extracellular Control of Gene Expression
Greg Somerville
Staphylococcus Aureus and S. Epidermidis Infections
Madhavan Soundararajan
Biochemical and Environmental Regulation of Carbon and Nitrogen Isotope Fractionation in Plants; Stomatal Biochemistry
Paul Staswick
Plant Molecular Biology; Crop Plant Quality; Disease Resistance Mechanisms
Julie Stone
Plant Molecular Biology; Programmed Cell Death; Signal Transduction
Karin van Dijk
Host/pathogen Interactions; Algal Biology
Mark Wilson
Protein X-ray Crystallography; Neurodegenerative Diseases
Charles Wood
Molecular Biology of HIV and Human Herpesviruses
Janos Zempleni
Chromatin Biology; Genome Stability; (Stem) Cell Differentiation; Adipocytes; Water-Soluble Vitamins; Gene Regulation
Jing Zhang
Endocytic Trafficking of Fatty Acids using Saccharomyces Cerevisiae as an Experimental Model
Limei Zhang
Structure-Function Relationships of Metallomolecules Involved in Redox Sensing, Metal Homeostasis and Detoxification, Metalloproteins Involved in Biotic and Abiotic Stress Sensing or Detoxification

Departments: Have an update for this page? Contact Stacy Dam.

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