Graduate Degree Program Summary

Graduate programs offered

Earn a Graduate Degree

  • MS in Biochemistry (30-36 cr, Option I, II, or III) with optional specialization:
    • Bioinformatics
    • Molecular Mechanisms of Disease
  • PhD in Biochemistry (90 cr) with optional specialization:
    • Bioinformatics
    • Molecular Mechanisms of Disease
Areas of Study

These informal areas of focus may help to shape your course of study but they will not appear on transcripts.

  • Structural and Chemical Basis of Protein Function
  • Metabolism and Metabolic Engineering
  • Molecular Mechanisms of Disease
  • Plant and Microbial Biochemistry and Biotechnology

Online and Distance Opportunities

Some online coursework may be available for your program; contact dept. for details.

Contacts for Biochemistry

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Other Ways to Connect

On the Web

Graduate Chair
Dr. Mark Wilson

Graduate Admissions Co-Chair
Dr. Ed Harris

Graduate Admissions Co-Chair
Dr. Limei Zhang

Campus Address
N200 Beadle Center
Lincoln NE 68588-0664

Promo image for Biochemistry

Application checklist and deadlines

1. Required by Graduate Studies

2. Required by Biochemistry

  • Minimum English proficiency: Paper TOEFL 550, Internet TOEFL 79, IELTS 6.5
  • Three recommendation letters
  • Curriculum Vitae or Resume
  • List of published papers, if applicable
  • Statement of purpose

When sending GRE or TOEFL scores, our institution code is 6877 and a department code is not needed.

Application Deadlines for Biochemistry
  • December 15 for Fall. September 1 for Spring.

Application/admission is for entry in a specific term and year. Our academic year is divided into 3 terms: Fall (August-December), Spring (January-May), and Summer (multiple sessions May-August). Some programs accept new students only in certain terms and/or years.


The Center for Biological Chemistry (CBC) offers a graduate program which is interdisciplinary. Research opportunities exist in a variety of areas, including plant biochemistry, photosynthesis research, plant and animal metabolism, metallobiochemistry, enzyme mechanisms, cancer biology, redox biology, and molecular biology. Biochemistry faculty and participating faculty in animal science, agronomy, chemistry, and biological sciences contribute to the program.

Instructional programs are focused in laboratories and classrooms within the George W. Beadle Center for Genetics and Biomaterials Research. The Beadle Center houses most of the department-affiliated faculty members in state-of-the-art research laboratories and many of the Center for Biotechnology and Redox Biology Center core facilities. These facilities include mass spectrometry, spectroscopy, X-ray crystallography, plant transformation, and microscopy facilities. Also available are an outstanding full-service plant growth and greenhouse complex, and equipment for fluorescence-activated cell analysis, surface plasmon resonance, analytical ultracentrifugation and next-generation high-throughput DNA sequencing.

Courses and More

Faculty and research

Where available, names link to bios or homepages and contact card icons () link to directory listings with address, phone, and email.

Jiri Adamec

Oxidative Stress; Proteomics; Metabolomics; Cancer; Neurodegenerative Diseases; Predictive Biomarkers

Donald Becker

Redox Enzymology; Gene Regulation; Multifunctional Proteins; Proline Metabolism

Paul Black

Fatty Acid Transport; Fatty Acid Transport Proteins; Metabolic Transformations

Nicole Buan Murphy

Energy Conservation; Redox Biochemistry; Microbial Physiology

Edgar Cahoon

Functional Genomics; Sphingolipid Metabolism and Function; Nutritional Biofortification of Crop Plants

Concetta DiRusso

Nutrient Fatty Acids; Fatty Acid-Dependent Regulation; Fatty Acids in Health and Disease

Katarzyna Glowacka

My long-term research goal is to improve the resistance of crops to abiotic stresses, in particular to low water availability and chilling temperatures.

Mark Griep

Biodefense Research; Antibiotic Discovery; Pathogenic Bacteria; DNA Replication; Bioinformatics

Edward Harris

Stabilin Receptors; Glycosaminoglycan Metabolism; Receptor-Mediated Endocytosis

Tomas Helikar

Computational Modeling of Dynamics in Large-Scale Biological/Biochemical Networks; Host-Pathogen Networks; Development of Technologies to Enable an Integrated Systems Biology Approach in Research and the Classroom

Oleh Khalimonchuk

Redox Sensitive Signaling Systems

Jaekwon Lee

Metal-Ion Metabolism; Copper Transporters; Oxidative Stress Response

Jonathan Markham

Sphingolipid Metabolism in Plants and Algae

Kenneth Nickerson

Fungal Dimorphism; Quorum Sensing in Fungi; Bacterial Detergent Resistance; Microbial Insecticides

Toshihiro Obata

Seung-Hyun Ro

Adipocyte Biology; Autophagy/Mitophagy; Mitochondria Metabolism; Oxidative Stress; Sestrins/mTOR Signaling; Nutrient Sensing and Metabolic Signaling in Obesity and Cancer

Rebecca Roston

Role of Membrane Dynamics in the Plant Cell using Combination of Genetics, Molecular Biology, Protein Biochemistry, and Biophysical Approaches; Membrane Response to Freezing Stress; Understanding Molecular Mechanisms of Photosynthetic Membrane Biogenesis

Erin Sayer

Madhavan Soundararajan

Biochemical and Environmental Regulation of Carbon and Nitrogen Isotope Fractionation in Plants; Stomatal Biochemistry

Julie Stone

Plant Molecular Biology; Programmed Cell Death; Signal Transduction

Xinghui Sun

Long Non-coding RNA Research in Metabolic Disease

Alex Vecchio

Mark Wilson

Protein X-ray Crystallography; Neurodegenerative Diseases

Charles Wood

Molecular Biology of HIV and Human Herpesviruses

Jing Zhang

Endocytic Trafficking of Fatty Acids using Saccharomyces Cerevisiae as an Experimental Model

Limei Zhang

Structure-Function Relationships of Metallomolecules Involved in Redox Sensing, Metal Homeostasis and Detoxification, Metalloproteins Involved in Biotic and Abiotic Stress Sensing or Detoxification

Karin van Dijk

Host/pathogen Interactions; Algal Biology

This summary page is maintained by Graduate Studies.
For additional details check out the dept./program website: Biochemistry.

Departments: Have an update for this summary? Contact Kurt Mueller.